Environmental change has profound impacts on ecological communities. It not only alters species composition but has long-lasting effects on more subtle aspects of biodiversity. Aside from species loss it can also cause the extinction and re-structuring of ecological interactions and the networks that they form. This ‘architecture of biodiversity’ is fundamental for the maintenance of ecosystem organisation, function and stability. Hence, if we are to predict the impacts of environmental change on ecosystems, it is of utmost importance that we understand the effects of its different drivers on the structure of networks of ecological interactions. In this lecture, I will introduce the concepts of climate warming and habitat loss as main environmental drivers of change on natural communities, with a particular focus on how they impact species interaction networks. We will look at the different mechanisms through which these drivers of change can affect the shape of ecological networks and how they can be understood through the lens of ecological theory. By combining these effects I will summarise a general framework to better understand and predict changes in ecological networks under environmental change.